Discovery of new species of mesoparasitic pennellid (Copepoda: Siphonostomatoida) from the endemic mesopelagic lightfish Vinciguerria mabahiss in the Red Sea

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Table 1

List of valid species of Cardiodectes Wilson C.B., 1917, including their geographic distribution and host fish species.

Cardiodectes species	Host family	Host species	Locality	Region	Reference(s)
C. anchorellae ^†	Engraulidae	Stolephorus tri	India (Madras, Kerala)	Indian Ocean	[8, 13, 35]
	Engraulidae	Thryssa hamiltonii	Sri Lanka	Indian Ocean	[24, 35]
	Engraulidae	Stolephorus indicus	India (Madras, Kerala)	Indian Ocean	[13]
C. bellottii ^†	Myctophidae	Benthosema glaciale	Mediterranean Sea	Mediterranean	[16, 22, 54]
	Myctophidae	Ceratoscopelus townsendi	Catalina Basin, California	Pacific Ocean	[32]
	Myctophidae	Ceratoscopelus warmingii	Eiao Island, Marquesas Archipelago	Pacific Ocean	[5]
	Myctophidae	Diaphus rafinesquii	Mediterranean Sea	Mediterranean	[22]
	Myctophidae	Diaphus suborbitalis	Japan (Misaki, Owase)	Pacific Ocean	[43, 54]
	Myctophidae	Diaphus theta	Catalina, Oregon Coast, San Clemente, Santa Barbara, Santa Cruz, San Pedro, Catalina Basins, California	Pacific Ocean	[30–32, 40]
	Myctophidae	Gonichthys cocco	Cape Point, South Africa; Off Mauritania, West Africa	Atlantic Ocean	[3, 32]
	Myctophidae	Hygophum benoiti	Mediterranean Sea	Mediterranean	[22]
	Myctophidae	Lampadena dea	Eiao Island, Marquesas Archipelago	Pacific Ocean	[5]
	Myctophidae	Lampanyctodes hectoris	South Africa (Cape Point, Saldanha Bay)	Atlantic Ocean	[3, 45]
	Myctophidae	Lampanyctus ritteri	Catalina Basin, San Clemente, San Pedro Basin, California	Pacific Ocean	[31, 32]
	Myctophidae	Myctophum affine	Mediterranean	Mediterranean	[7]
	Myctophidae	Myctophum nitidulum	East coast of Hamahiga Island, Japan	Pacific Ocean	[49]
	Myctophidae	Myctophum sp.	São Tiago Island, Cape Verde Islands	Atlantic Ocean	[9]
	Myctophidae	Notoscopelus caudispinosus	Genoa, Italy	Mediterranean	[6]
	Myctophidae	Parvilux ingens	Catalina Basin, California	Pacific Ocean	[32]
	Myctophidae	Stenobrachius leucopsarus	Los Angeles, Oregon Coast, San Clemente, San Diego, San Pedro, Santa Barbara, Santa Cruz, Catalina Basins, California	Pacific Ocean	[15, 29–32, 53, 54]
	Myctophidae	Symbolophorus californiensis	Catalina, San Pedro, San Clemente Basins, California	Pacific Ocean	[32]
	Myctophidae	Tarletonbeania crenularis	San Clemente, San Pedro, Catalina Basins, California	Pacific Ocean	[32, 40]
	Sternoptychidae	Maurolicus muelleri	Mauritius	Indian Ocean	[33]
C. cristatus ^†	Myctophidae	Diaphus suborbitalis	Off the coast of Owase, Japan	Pacific Ocean	[43]
C. frondosus ^†	Myctophidae	Dasyscopelus spinosus	Nuku Hiva, Marquesas Archipelago	Pacific Ocean	[41]
C. longicervicus ^†	Myctophidae	Dasyscopelus asper	Off the coast of Owase, Japan	Pacific Ocean	[43]
C. asper ^§	Gobiidae	Trimma grammistes	Izu-Oshima Island, Japan	Pacific Ocean	[50]
C. bellwoodi ^§	Gobiidae	Istigobius nigroocellatus	Great Barrier Reef, Australia	Pacific Ocean	[49]
C. bertrandi ^§	Gobiidae	Eviota sp.	Loyalty Islands, New Caledonia	Pacific Ocean	[50]
C. boxshalli ^§	Scaridae	Nicholsina usta	Jamaica	Caribbean Sea	[4]
C. hardenbergi ^§	Engraulidae	Stolephorus spp.	Java Sea	Indian Ocean	[28]
C. krishnai ^§	Phosichthyidae	Vinciguerria lucetia	Arabian Sea	Arabian Sea	[35, 42]
C. roatanensis ^§	Scaridae	Nicholsina usta	Roatan Island, Honduras	Caribbean Sea	[46]
C. rotundicaudatus ^§	Gobiidae	Suruga fundicola	Sagami Bay, Japan	Pacific Ocean	[21]
C. rubosus ^§	Apogonidae	Apogon sp.	Salomakië Island, Alaska	Pacific Ocean	[26]
C. rubosus ^§	Dorosomatidae	Harengula clupeola	Cartagena, Columbia	Caribbean Sea	[52]
C. shini ^§	Gobiidae	Eviota sp.	Okinawa-jima Island, Ryukyu Islands, Japan	Pacific Ocean, East China Sea	[14, 49]
	Gobiidae	Eviota sebreei
	Gobiidae	Pleurosicya micheli
	Gobiidae	Priolepis sp.
C. spiralis ^§	Anthiadinae	Pseudanthias tuka	Massas Island, Papua New Guinea	Pacific Ocean	[4]
Cardiodectes sp.	Pinguipedidae	Parapercis sexfasciata	Off the Tanegashima, Japan	Pacific Ocean	[57]
C. vampire ^§	Chlorophthalmidae	Chlorophthalmus corniger	Arabian Sea	Arabian Sea	[2]
C. tofaili n. sp.^§	Phosichthyidae	Vinciguerria mabahiss	Red Sea	Red Sea	Present study

^{† “}Medusaeus” group, ^§ “Rubosus” group.

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