Truly a hyperparasite, or simply an epibiont on a parasite? The case of Cyclocotyla bellones (Monogenea, Diclidophoridae)

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Table 5

Combinations, hosts and localities of Cyclocotyla spp. in the literature. The current combination is in bold.

Parasites	Synonyms	Type host	Type locality	Source
*Cyclocotyla bellones*	Cyclobothrium charcoti	Belone belone	Italy, Mediterranean Sea	[72]
Otto, 1823	Dollfus, 1922			[35]
Cyclocotyla charcoti	Cyclobothrium charcoti ** Dollfus, 1922	Ceratothoa oestroides (female), buccal cavity of Trachurus trachurus	Spain, Atlantic Ocean	[34, 35, 60, 100]
(Dollfus, 1922) Price, 1943 **	Choricotyle charcoti * (Dollfus, 1922) Llewellyn, 1941 Cyclocotyla bellones
	*Allodiclidophora charcoti* (Dollfus, 1922) Yamaguti 1963
Cyclocotyla elongata (Goto, 1894) Price, 1943*	Diclidophora elongate Goto, 1894**	Buccal cavity of Dentex tumifrons, Cymothoa parasitic in this cavity	Japan, Pacific Ocean	[45, 60]
Cyclocotyla elongata (Goto, 1894) Price, 1943*	*Choricotyle elongata* (Goto, 1894) Llewellyn, 1941		Japan, Pacific Ocean	[45, 60]
Cyclocotyla smaris ** (Ijima, in Goto, 1894) Price, 1943	Diclidophora smaris *Ijima, in Goto, 1894 Octobothrium smaris* Ijima, in Goto, 1894 Choricotyle smaris (Ijima, in Goto, 1894) Llewellyn, 1941	Cymothoa of buccal cavity of Spicara smaris	Italy, Mediterranean Sea	[45, 60, 78]
Cyclocotyla hysteroncha Fujii, 1944*	*Choricotyle hysteroncha* (Fujii, 1944) Sproston, 1946	Gills of Haemulon striatum	USA, Atlantic Ocean	[43, 91]
Cyclocotyla labracis (Cerfontaine, 1895) Price, 1943	*Choricotyle labracis* (Cerfontaine, 1895) Llewellyn, 1941	Gills of Dicentrarchus labrax	North Sea, Atlantic Ocean	[19, 60, 78]
Cyclocotyla louisianensis * (Hargis, 1955)	*Hargicotyle louisianensis* (Hargis, 1955) Mamaev, 1972	Gills of Menticirrhus americanus	USA, Atlantic Ocean	[47, 64]
Cyclocotyla multaetesticulae * Chauhan, 1945	*Choricotyle multaetesticulae* (Chauhan, 1945) Sproston, 1946	Gills of a marine fish Pellona sp.	India, Indian Ocean	[25, 91]
Cyclocotyla neomaenis*	*Echinopelma neomaenis* (MacCallum, 1917) Raecke, 1945	Gills of Lutjanus analis	USA, Atlantic Ocean	[63, 81]
(MacCallum, 1917) Price, 1943	*Echinopelma neomaenis* (MacCallum, 1917) Raecke, 1945	Gills of Lutjanus analis	USA, Atlantic Ocean	[63, 81]
Cyclocotyla prionoti * (MacCallum, 1917) Price, 1943	*Orbocotyle prionoti* (MacCallum, 1917) Euzet & Suriano, 1975	Gills of Prionotus carolinus	USA, Atlantic Ocean	[78]
Cyclocotyla prionoti * (MacCallum, 1917) Price, 1943	Diclidophora neomaenis *MacCallum, 1917 Choricotyle neomaenis* (MacCallum,1917) Llewellyn, 1941 **	Gills of Prionotus carolinus	USA, Atlantic Ocean	[78]
Cyclocotyla squillarum **	Mesocotyle squillarum Parona & Perugia, 1889	Ovigerous lamellae of Bopyrus squillarum	Italy, Mediterranean Sea	[76]
(Parona & Perugia, 1889)	*Allodiclidophora squillarum* (Parona & Perugia, 1889)

Junior synonym.

^**

Senior synonym.

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