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Cited article:

Phylogeny, ontogeny and adaptive radiation in the superfamily Tydeoidea (Acari: Actinedida), with a reappraisal of morphological characters

HENRI M. ANDRÉ and ALEX FAIN
Zoological Journal of the Linnean Society 130 (3) 405 (2000)
https://doi.org/10.1111/j.1096-3642.2000.tb01636.x