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Morphometrics of Quadriacanthus spp. proposed by Gussev (1962) and modified by N’Douba et al. (1999). (A) Anchor: (a) length, (ab) base width, (e) point length; (Ap) Accessory piece length; (MCO) Male copulatory organ length; (C) Cuneus: (j) length, (i) width; (DB) Dorsal bar: (ct) center length, (h) median process length, (w) width, (x) length; (VB) Ventral bar: (w) width, (x) length; (H) Hook length; (Vg) Vaginal length; scale 20 µm.

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