Phylogenetic relationships by maximum likelihood (ML) analysis among sequences of mitochondrial 12S ribosomal RNA (A), mitochondrial cytochrome c oxidase subunit 1 (B) genes, and the internal transcribed spacer 1-5.8S ribosomal RNA region (C). The ML tree was derived from a general time-reversible model using a discrete gamma distribution (+G) with five rate categories and invariant sites (+I). Significant bootstrap support for the ML analysis with 1000 replicates and Bayesian analysis (BI) are shown above the nodes in the order ML/BI. An asterisk indicates <50% support for a node. The scale bar represents the genetic distance in single nucleotide substitutions. GenBank accession numbers are given within parentheses.